Cell

The molecular mechanism underlying the retention of intron-containing mRNAs in the nucleus is not understood. Here, we show that retention of intron-containing mRNAs in yeast is mediated by perinuclearly located Mlp1. Deletion of MLP1 impairs retention while having no effect on mRNA splicing. The Mlp1-dependent leakage of intron-containing RNAs is increased in presence of ts-prp18 delta, a splicing mutant. When overall pre-mRNA levels are increased by deletion of RRP6, a nuclear exosome component, MLP1 deletion augments leakage of only the intron-containing portion of mRNAs. Our data suggest, moreover, that Mlp1-dependent retention is mediated via the 5' splice site. Intriguingly, we found Mlp-proteins to be present only on sections of the NE adjacent to chromatin. We propose that at this confined site the perinuclear Mlp1 implements a quality control step prior to export, physically retaining faulty pre-mRNAs.

Source:http://purl.uniprot.org/citations/14718167

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http://purl.uniprot.org/cit...rdfs:commentThe molecular mechanism underlying the retention of intron-containing mRNAs in the nucleus is not understood. Here, we show that retention of intron-containing mRNAs in yeast is mediated by perinuclearly located Mlp1. Deletion of MLP1 impairs retention while having no effect on mRNA splicing. The Mlp1-dependent leakage of intron-containing RNAs is increased in presence of ts-prp18 delta, a splicing mutant. When overall pre-mRNA levels are increased by deletion of RRP6, a nuclear exosome component, MLP1 deletion augments leakage of only the intron-containing portion of mRNAs. Our data suggest, moreover, that Mlp1-dependent retention is mediated via the 5' splice site. Intriguingly, we found Mlp-proteins to be present only on sections of the NE adjacent to chromatin. We propose that at this confined site the perinuclear Mlp1 implements a quality control step prior to export, physically retaining faulty pre-mRNAs.lld:uniprot
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http://purl.uniprot.org/cit...uniprot:authorRomano A.lld:uniprot
http://purl.uniprot.org/cit...uniprot:authorNehrbass U.lld:uniprot
http://purl.uniprot.org/cit...uniprot:authorJacquier A.lld:uniprot
http://purl.uniprot.org/cit...uniprot:authorGadal O.lld:uniprot
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http://purl.uniprot.org/cit...uniprot:authorGaly V.lld:uniprot
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http://purl.uniprot.org/cit...uniprot:pages63-73lld:uniprot
http://purl.uniprot.org/cit...uniprot:titleNuclear retention of unspliced mRNAs in yeast is mediated by perinuclear Mlp1.lld:uniprot
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